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Research article · 2024 · Full text

International Journal of Research in Agronomy

Ichthyofaunal diversity of Milak, a small river of Nagaland

Nyamkham P PhomPampa BhattacharjeeDiamond Rajakumar Tenali

Volume 7(6S) · Pages 250–253 · DOI: 10.33545/2618060x.2024.v7.i6sd.891 ↗

Research areasFish Biology and Ecology StudiesFish Ecology and Management StudiesSouth Asian Studies and Conflicts

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Abstract

Author-written

The present study is targeted to investigate the fish diversity, their present IUCN conservation status, and economic value of Milak, a small river of Nagaland. Milak River originated in Langpangkong of Mokokchung and is effectively marked by the moderate and progressing flow towards the plains. Tuli is one of the most populous towns that are located near this river district in Nagaland. Tsurong is a major tributary of the Milak River.

Summary

Source-checked

Plain-language reading of the full paper, prepared with AI and checked against the text. Each point links to the authors' own words.

  1. Why it matters

    • Nagaland is a small hilly state in northeastern India covering 16,579 sq. km and is located within the Indo-Burma Biodiversity Hotspot. View evidence ↗

      Why it matters

      Nagaland is a small hilly state in northeastern India covering 16,579 sq. km and is located within the Indo-Burma Biodiversity Hotspot.

      In the paper's own words

      “Nagaland, a small hilly state (area of 16,579 sq. km) in northeastern India, is a part of the Indo-Burma Biodiversity Hotspot.”

      Page 1

      Found in: Introduction

      Read it in the full textOpen PDF at page 1 ↗
    • Fish are significant biological resources used for food, the aquarium trade, and the sports industry. View evidence ↗

      Why it matters

      Fish are significant biological resources used for food, the aquarium trade, and the sports industry.

      In the paper's own words

      “Fish is one of the important bioresources which can be used as food, in Aquarium fish, and sports industry (1).”

      Page 1

      Found in: Introduction

      Read it in the full textOpen PDF at page 1 ↗
    • Freshwater ecosystems are among the most endangered in the world, despite covering only about 0.01% of the Earth's surface water. View evidence ↗

      Why it matters

      Freshwater ecosystems are among the most endangered in the world, despite covering only about 0.01% of the Earth's surface water.

      In the paper's own words

      “Freshwater ecosystems contain only around 0.01% of the world’s surface water cover; meanwhile, freshwater ecosystems have become the most endangered ecosystems in the world”

      Page 2

      Found in: Results And Discussion

      Read it in the full textOpen PDF at page 2 ↗
  2. The question

    • The study aimed to investigate the fish diversity, economic value, and IUCN conservation status of species in the Milak River of Nagaland. View evidence ↗

      The question

      The study aimed to investigate the fish diversity, economic value, and IUCN conservation status of species in the Milak River of Nagaland.

      In the paper's own words

      “The present study is targeted to investigate the fish diversity, their present IUCN conservation status, and economic value of Milak, a small river of Nagaland.”

      Page 1

      Found in: Materials And Methods

      Read it in the full textOpen PDF at page 1 ↗
  3. What they did

    • The survey was conducted in the Milak River of the Mokokchung district of Nagaland during the winter and pre-monsoon seasons. View evidence ↗

      What they did

      The survey was conducted in the Milak River of the Mokokchung district of Nagaland during the winter and pre-monsoon seasons.

      In the paper's own words

      “The survey was carried out at of Milak River of Mokokchung district of Nagaland (Fig: 1). The study period was during the pre-monsoon and winter seasons.”

      Page 1

      Found in: Materials And Methods

      Read it in the full textOpen PDF at page 1 ↗
    • Fish specimens were collected using cast nets with the assistance of local artisanal fishers. View evidence ↗

      What they did

      Fish specimens were collected using cast nets with the assistance of local artisanal fishers.

      In the paper's own words

      “Specimens were collected with the help of a cast net along with local artisanal fishers.”

      Page 1

      Found in: Materials And Methods

      Read it in the full textOpen PDF at page 1 ↗
    • Specimens were preserved in 10% formalin, and measurements were recorded to 0.1 mm using digital calipers. View evidence ↗

      What they did

      Specimens were preserved in 10% formalin, and measurements were recorded to 0.1 mm using digital calipers.

      In the paper's own words

      “The collected specimens were preserved in 10% formalin for further analysis. The descriptions are based on formalin-preserved specimens. Measurements were taken point to point with digital calipers to 0.1 mm.”

      Page 1

      Found in: Materials And Methods

      Read it in the full textOpen PDF at page 1 ↗
    • The conservation status of the identified fish species was determined using the IUCN Red List. View evidence ↗

      What they did

      The conservation status of the identified fish species was determined using the IUCN Red List.

      In the paper's own words

      “The conservation status of the fish species the conservation status of the fish species is based on the IUCN (www.iucnredlist.org.in).”

      Page 1

      Found in: Materials And Methods

      Read it in the full textOpen PDF at page 1 ↗
  4. What they found

    • The study identified a total of 20 fish species belonging to five families: Cyprinidae, Sisoridae, Channidae, Mastacembelidae, and Badidae. View evidence ↗

      What they found

      The study identified a total of 20 fish species belonging to five families: Cyprinidae, Sisoridae, Channidae, Mastacembelidae, and Badidae.

      In the paper's own words

      “A total of 20 species from the families Cyprinidae, Sisoridae, Channidae, Mastacembelidae, and Badidae were collected.”

      Page 2

      Found in: Results And Discussion

      Read it in the full textOpen PDF at page 2 ↗
    • The species Olyra kempi was listed as Data Deficient (DD). View evidence ↗

      What they found

      The species Olyra kempi was listed as Data Deficient (DD).

      In the paper's own words

      “12 | Siluriformes | Bagridae | Olyra kempi | F | DD”

      Page 2

      Found in: Table 1

      Read it in the full textOpen PDF at page 2 ↗
    • Danio aequipinnatus was recorded as Not Evaluated (NE) for its conservation status. View evidence ↗

      What they found

      Danio aequipinnatus was recorded as Not Evaluated (NE) for its conservation status.

      In the paper's own words

      “3 | Cypriniformes | Cyprinidae | Danio aequipinnatus | F, O | NE”

      Page 2

      Found in: Table 1

      Read it in the full textOpen PDF at page 2 ↗
    • The authors noted that two species within the genera Garra and Badis appear to be new to science, though this requires further research for confirmation. View evidence ↗

      What they found

      The authors noted that two species within the genera Garra and Badis appear to be new to science, though this requires further research for confirmation.

      In the paper's own words

      “revealing the existence of some ambiguous fish species that seem to be new to science.”

      Page 2

      “Among the Genus Garr sp. and Badis sp. Two seems to be new species, further research work required for confirmation.”

      Page 4

      Found in: Results And Discussion

      Read it in the full textOpen PDF at page 2 ↗
  5. What it means

    • The authors recommend immediate fish conservation measures and more frequent exploratory surveys to document diversity before species go extinct. View evidence ↗

      What it means

      The authors recommend immediate fish conservation measures and more frequent exploratory surveys to document diversity before species go extinct.

      In the paper's own words

      “Suggesting more frequent fish exploratory surveys to know the valuable fish diversity before extinction and the need of immediate fish conservation implementations.”

      Page 2

      Found in: Results And Discussion

      Read it in the full textOpen PDF at page 2 ↗
    • Further research is required to confirm whether the identified specimens of Garra sp. and Badis sp. are indeed new species. View evidence ↗

      What it means

      Further research is required to confirm whether the identified specimens of Garra sp. and Badis sp. are indeed new species.

      In the paper's own words

      “Among the Genus Garr sp. and Badis sp. Two seems to be new species, further research work required for confirmation.”

      Page 4

      Found in: Conclusions

      Read it in the full textOpen PDF at page 4 ↗

Figures & tables

From the original paper
Showing the IUCN dataTable 1Page 2One of the sampling locations of Milak RiverFigure 1Page 3One of the sampling locations of Milak RiverFigure 1Page 3One of the sampling locations of Milak RiverFigure 1Page 3Some collected species from Milak riverFigure 2Page 3Some collected species from Milak riverFigure 2Page 3Percentage of fishes according to their importance in FisheriesFigure 3Page 4
FULL TEXT Original article · reflowed for reading

Introduction

Nagaland, a small hilly state (area of 16,579 sq. km) in northeastern India, is a part of the Indo- Burma Biodiversity Hotspot. The state is bordered by Arunachal Pradesh to the North, Manipur to the South, and Myanmar and Assam to the east and west, respectively. The diversity of topographic and climatic conditions has made the state rich with various flora and fauna. Fish is one of the important bioresources which can be used as food, in Aquarium fish, and sports industry (1).

In the present study, we provide a preliminary checklist of the fish fauna of Milak River. A rigorous collection of fish from the river obtained a good number of species diversity as the area lies in a hotspot.

Materials And Methods

Study site: The survey was carried out at of Milak River of Mokokchung district of Nagaland (Fig: 1). The study period was during the pre-monsoon and winter seasons. Specimens were collected with the help of a cast net along with local artisanal fishers. All the essential data such as place of collection, number of fish, body color, and body marking were recorded.

The collected specimens were preserved in 10% formalin for further analysis. The descriptions are based on formalin-preserved specimens. Measurements were taken point to point with digital calipers to 0.1 mm. For identification and classification, (2) and (3) were followed.

The conservation status of the fish species the conservation status of the fish species is based on the IUCN (www.iucnredlist.org.in). Table 1 shows the list of fish recorded from Milak River along with their economic value and the IUCN 2024 status. https://www.agronomyjournals.com Table 1: Showing the IUCN data Sl. No. Order Family Species Economic value IUCN-2024 Status 1 Cypriniformes Cyprinidae Barilius barna F; O LC 2 Cypriniformes Cyprinidae Opsarius bendelesis F; O LC 3 Cypriniformes Cyprinidae Danio aequipinnatus F, O NE 4 Cypriniformes Cyprinidae Danio dangila F LC 5 Cypriniformes Cyprinidae Devario devario F LC 6 Cypriniformes Cyprinidae Pethia ticto F; O LC 7 Cypriniformes Cyprinidae Tor tor S LC 8 Cypriniformes Cyprinidae Schizothorax richardsonii F LC 9 Cypriniformes Cyprinidae Garra kempi F; S LC 10 Anabantiformes Badidae Badis badis F, O LC 11 Siluriformes Bagridae Olyra longicaudatus F; O LC 12 Siluriformes Bagridae Olyra kempi F DD 13 Siluriformes Bagridae Sperata seenghala F VU 14 Siluriformes Sisoridae Pseudecheneis sirenica O VU 15 Siluriformes Sisoridae Glyptothorax cavia O LC 16 Siluriformes Sisoridae Glyptothorax indicus O LC 17 Siluriformes Sisoridae Exostoma labiatum O LC 18 Siluriformes Sisoridae Exostoma berdmorei O LC 19 Synbranchiformes Mastacembelidae Macrognathus pancalus F; O LC 20 Anabantiformes Channidae Channa punctata F; O LC DD- Data Deficient, LC-Least concern, NE- Not evaluated, VU- Vulnerable, F-Food, O-Ornamental, S-Sport

Table 1. Showing the IUCN data
Sl. No.OrderFamilySpeciesEconomic valueIUCN-2024 Status
1CypriniformesCyprinidaeBarilius barnaF; OLC
2CypriniformesCyprinidaeOpsarius bendelesisF; OLC
3CypriniformesCyprinidaeDanio aequipinnatusF, ONE
4CypriniformesCyprinidaeDanio dangilaFLC
5CypriniformesCyprinidaeDevario devarioFLC
6CypriniformesCyprinidaePethia tictoF; OLC
7CypriniformesCyprinidaeTor torSLC
8CypriniformesCyprinidaeSchizothorax richardsoniiFLC
9CypriniformesCyprinidaeGarra kempiF; SLC
10AnabantiformesBadidaeBadis badisF, OLC
11SiluriformesBagridaeOlyra longicaudatusF; OLC
12SiluriformesBagridaeOlyra kempiFDD
13SiluriformesBagridaeSperata seenghalaFVU
14SiluriformesSisoridaePseudecheneis sirenicaOVU
15SiluriformesSisoridaeGlyptothorax caviaOLC
16SiluriformesSisoridaeGlyptothorax indicusOLC
17SiluriformesSisoridaeExostoma labiatumOLC
18SiluriformesSisoridaeExostoma berdmoreiOLC
19SynbranchiformesMastacembelidaeMacrognathus pancalusF; OLC
20AnabantiformesChannidaeChanna punctataF; OLC

Table text extracted from the original PDF; superscript letters are shown inline. The original table image is below.

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Showing the IUCN data↗ Enlarge
Table 1
Showing the IUCN data

Showing the IUCN data

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Original paper tableSource PDF · page 2 ↗

Results And Discussion

Freshwater ecosystems contain only around 0.01% of the world’s surface water cover; meanwhile, freshwater ecosystems have become the most endangered ecosystems in the world, and the biodiversity that they support is vulnerable due to human activities and environmental changes. However, the Eastern Himalaya region has been identified as a freshwater biodiversity hotspot (4) and (5). The current study is the result of an exploration of fish diversity of Milak, a small river in Nagaland. A total of 20 species from the families Cyprinidae, Sisoridae, Channidae, Mastacembelidae, and Badidae were collected.

A greater number of species of the genera Danio, Opsarius, Devario, Badis, Schizothorax, Exostoma, and Glyptothorax were collected in present fish explorations. Due to the importance of taxonomical confirmation of species, the present study was conducted in a small river named Milak of the Brahmaputra basin revealing the existence of some ambiguous fish species that seem to be new to science. The study reveals the presence of twenty species belonging to 4 orders, 7 families and 16 genera. Cypriniformes dominated by 9 species followed by 8 siluriformes and each one of anabantiformes and synbranchiformes.

A greater number of species of the genera Garra, Opsarius, Psilorhynchus, Devario, Badis, Schizothorax, Exostoma, Garra and Glyptothorax (Fig-2) were recorded abundantly from the river. Suggesting more frequent fish exploratory surveys to know the valuable fish diversity before extinction and the need of immediate fish conservation implementations. https://www.agronomyjournals.com Fig 1: One of the sampling locations of Milak River Fig 2: Some collected species from Milak river https://www.agronomyjournals.com Fig 3: Percentage of fishes according to their importance in Fisheries

Figure 3. Percentage of fishes according to their importance in Fisheries
Percentage of fishes according to their importance in Fisheries↗ Enlarge
Figure 3
Percentage of fishes according to their importance in Fisheries

Percentage of fishes according to their importance in Fisheries

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Original paper figureSource PDF · page 4 ↗
Figure 2. Some collected species from Milak river
Some collected species from Milak river↗ Enlarge
Figure 2
Some collected species from Milak river

Some collected species from Milak river

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Original paper figureSource PDF · page 3 ↗
Figure 2. Some collected species from Milak river
Some collected species from Milak river↗ Enlarge
Figure 2
Some collected species from Milak river

Some collected species from Milak river

Open full-size image ↗
Original paper figureSource PDF · page 3 ↗
Figure 1. One of the sampling locations of Milak River
One of the sampling locations of Milak River↗ Enlarge
Figure 1
One of the sampling locations of Milak River

One of the sampling locations of Milak River

Open full-size image ↗
Original paper figureSource PDF · page 3 ↗
Figure 1. One of the sampling locations of Milak River
One of the sampling locations of Milak River↗ Enlarge
Figure 1
One of the sampling locations of Milak River

One of the sampling locations of Milak River

Open full-size image ↗
Original paper figureSource PDF · page 3 ↗
Figure 1. One of the sampling locations of Milak River
One of the sampling locations of Milak River↗ Enlarge
Figure 1
One of the sampling locations of Milak River

One of the sampling locations of Milak River

Open full-size image ↗
Original paper figureSource PDF · page 3 ↗

Conclusions

Total 20 Fish species Identified and recorded from Milak river of Nagaland. Among the Genus Garr sp. and Badis sp. Two seems to be new species, further research work required for confirmation. Suggesting more frequent fish exploratory surveys to know the valuable fish diversity before extinction and the need of immediate fish conservation implementations.

Acknowledgments

The authors extend their thankfulness to the Vice Chancellor, Central Agricultural University, Imphal, India and the Dean, College of Fisheries, Central Agricultural University, Tripura for providing the necessary facilities for carrying out the study.

References

5 in the original paper
  1. Gurumayum SD, Kosygin L, Tamang L. Ichthyofaunal diversity of Arunachal Pradesh, India: A part of Himalaya biodiversity hotspot. Int J Fish Aquat Stud. 2016;4(2):337- 346.
  2. Nath P, Dey SC. Fish and Fisheries of Northeastern India. Vol. I: Arunachal Pradesh. 1st ed.; c2000. p. 217.
  3. Vishwanath W. Freshwater fishes of the Eastern Himalayas. Elsevier Academic Press; c2021. p. 431.
  4. Das BK, Boruah P, Kar D. Fish diversity and drainage analysis of River Siang, East Siang district of Arunachal Pradesh. Biosci Discov. 2014;6(1-I):16-20.
  5. IUCN. IUCN Red List of Threatened Species. Version 2023-2024. [Internet]. Available from: <www.iucn.redlist.org>. Accessed 2024.